School of Biology
University of Leeds
Leeds, LS2 3JT

Lab: +44 (0113) 343 5578
Fax: +44 (0113) 343 2835

2012-present    Lecturer in Animal Biology (University of Leeds)
2010–2012       Ontario MRI Postdoctoral Fellow (Carleton University)
2009–2010       Government of Canada Postdoctoral Fellow (Carleton University)

2005–2009       PhD “The Impacts of Environmental Warming on the Biology of British Odonata” (University of Liverpool)
2002–2005       BSc (Hons) Zoology (1st Class) (University of Liverpool)

Studentships and Fellowships

  • 2015 Marie Curie Fellowship (€230k)
  • 2010 Ontario MRI Postdoctoral Fellowship ($10,000 research funding)
  • 2009 Government of Canada Postdoctoral Fellowship
  • 2009 Natural Environment Research Council Policy Secondment to Welsh Government
  • 2005 Natural Environment Research Council Studentship ($21,700 research funding)


  • 2014 Royal Society Research Grant (£15k)
  • 2011 Freshwater Biological Association Hugh Carey Gilson Award ($5,780)
  • 2010 British Ecological Society Small Ecological Project Grant ($3,870)
  • 2008 Royal Entomological Society Conference Participation Grant ($400)


  • 2009 Royal Entomological Society Student Award
  • 2005 Liverpool Biological Society Prize in Zoology
  • 2004 Agnes Sinclair Undergraduate University Scholarship
  • 2003 Agnes Sinclair Undergraduate University Scholarship

I referee papers for (in no particular order, and probably missing a few…) Ecological Entomology, Freshwater Biology, Global Change Biology, Central European Journal of Biology, Journal of Zoology, International Journal of Biometeorology, PLoS ONE, Biological Journal of the Linnaean Society, Insect Conservation and Diversity, Ecology Letters, American Naturalist, Evolutionary Ecology, Hydrobiologia, Journal of Insect Science, and Oecologia.

I am an Associate Editor of Insect Conservation and Diversity and am on the Editorial Board of Ecological Entomology.


October 2014 Invited seminar (University of Hull, UK)
Title: “The evolution of imperfect mimicry”

September 2014 IALE UK Annual Meeting (London, UK)
Oral Presentation: “A comparative analysis of biodiversity in urban ponds in the UK”

July 2014 Humberhead Levels Monitoring Meeting (Doncaster, UK)
Invited talk: “Capture-mark-recapture (CMR) as a method for species monitoring at a landscape scale”

July 2014 European Congress on Odonatology (Montpellier, France)
Oral presentation: “Genetic and morphological variation at an expanding range margin in a damselfly”

June 2014 Invited seminar (University of Leuven, Belgium)
Title: “Ecological and evolutionary drivers of macroecological patterns in Odonata”

January 2014 Invited seminar (Manchester Metropolitan University, UK)
Title: “Evolution of imperfect mimicry”

October 2013 Ecology of inland waters workshop (Donana National Park, Spain)
Oral presentation: “Urban ponds and aquatic biodiversity”

August 2013 International Congress on Ecology (London, UK)
Poster presentation: “Mimicry under climate change: potential for mimetic decoupling”

August 2013 International Congress on Ethology (Newcastle, UK)
Invited talk: “Evolution of imperfect mimicry”

July 2013 World Congress on Odonatology (Freising, Germany)
Oral presentation: “Odonata as barometers for global change”

March 2013 Invited seminar (University of York, UK)
Title: “Evolution of imperfect mimicry”

July 2012 Evolution 2012 (Ottawa, Canada)
Oral presentation: “The evolution of imperfect mimicry”

May 2011 Canadian Society for Ecology and Evolution (Banff, Canada)
Oral presentation: “Evidence for convergent character displacement in a damselfly”

May 2010 Canadian Society for Ecology and Evolution (Quebec City, Canada)
Oral presentation: “Present and future diversity of North American Odonata”

Aug 2009 International Congress on Ecology (Brisbane, Australia)
Oral presentation: “Global change and the predatory niche in aquatic ecosystems”

Jan 2009 Wales Environment into Policy Forum (Cardiff, Wales)
Meeting for policy-makers and scientists on the environmental research in Wales

Oct 2008 Royal Entomological Society Climate Change SIG (York, England)
Poster presentation: “Contrasting patterns of genetic and morphometric variation in a damselfly undergoing rapid range expansion”

Sept 2008 Royal Entomological Society (University of Plymouth, England)
Oral presentation: “Connectivity of odonate habitat in north west England”.

May 2008 European Pond Conservation Network (Valencia, Spain)
Oral presentation: “Odonata and climate change in pond landscape of north west England”.

Sept 2007 British Ecological Society Annual Meeting (University of Glasgow, Scotland)
Poster: “Climate change impacts on the phenology of Odonata”.

Professional courses
June 2008 National Pond Survey Methods (Freshwater Biological Association, Cumbria)
Course on standardised methods used in surveying ponds.
Mar 2008 Occupancy 2008 (University of St. Andrews, UK)
Course on the statistical methods used in estimating animal distributions.
Nov 2006 Advances in Ecology (Imperial College, UK)
Course on ecological modelling using the R programming language.


Current teaching:
  • Research Experience and Skills Level 2 (BLGY2301)
  • Mediterranean Ecology Field Course (BLGY2313)
  • 20 Credit Research Projects (BLGY3021)
  • 40 Credit Research Projects (BLGY3340)
  • Advanced Topics in Ecology (BLGY3133)
  • Introduction to GIS Skills for Ecologists (BLGY5117M, Module Manager)
  • Masters Mediterranean Ecology Field Course (BLGY5156M)
  • Biodiversity and Conservation MRes Research Project 1 (BLGY5198M)
Previous teaching:
  • Research Experience and Skills Level 1 (BLGY1304, Leeds)
  • Insect Identification (BLGY5162M, Module Manager, Leeds)
  • Biodiversity and Conservation MSc and MRes Summer Project (BLGY5191M, Leeds)
  • Conservation Biology (BIOL3602, Carleton University)
  • Biology Honours Projects (BIOL4908, Carleton University)
  • Postgraduate teaching assistant (several courses, University of Liverpool)


  • British Ecological Society (member) 2006 – present
  • Freshwater Biological Association (member) 2007 – present
  • Royal Entomological Society (member) 2007 – 2013; (fellow) 2013 – present
  • European Pond Conservation Network (member) 2008 – present
  • Canadian Society for Ecology and Evolution (member) 2010 – present


Hassall, C. and Anderson, S.* (in press) Stormwater ponds can contain comparable biodiversity to unmanaged wetlands in urban areas, Hydrobiologia.

Noble, A.* and Hassall, C. (in press) Poor ecological quality of urban ponds in northern England: causes and consequences, Urban Ecosystems.

Fraser, D., Hassall, C., Gorelick, R., and Rybczynski, N. (2014) Mean annual precipitation explains spatiotemporal patterns of Cenozoic mammal beta diversity and latitudinal diversity gradients in North America. PLoS ONE, 9: e106499.

Hassall, C. (2014) Continental variation in wing pigmentation in Calopteryx damselflies is related to the presence of heterospecifics, PeerJ 2:e438

Hassall, C. (2014) The ecology and biodiversity of urban ponds, WIREs Water, 1: 187–206.

Penney, H.D., Hassall, C., Skevington, J.H., Lamborn, B., and Sherratt, T.N. (2014) The relationship between morphological and behavioral mimicry in hover flies (Diptera: Syrphidae), American Naturalist, 183: 281-289.

Easley, J.* & Hassall, C. (2014) Field estimates of survival do not reflect ratings of mimetic similarity in wasp-mimicking hoverflies, Evolutionary Ecology, 28, 387-396.

Hassall, C., Keat, S., Thompson, D.J., and Watts, P.C. (2014) Bergmann’s rule is maintained during a rapid range expansion in a damselfly, Global Change Biology, 20: 475–482.

Webster, R.J., Hassall, C., Herdman, C.M., Godin, J.-G.J. and Sherratt, T.N. (2013) Disruptive camouflage impairs object recognition, Biology Letters, 9: 20130501.

Hassall, C. (2013) “Going green”: self-archiving as a means for dissemination of research output in ecology and evolution, Ideas in Ecology and Evolution, 5: 93-98.

Hossie, T.H., Hassall, C., Knee, W., and Sherratt, T.N. (2013) Species with a chemical defense, but not chemical offense, live longer, Journal of Evolutionary Biology, 26: 1598-1602.

Hassall, C. (2013) Time stress and temperature explain continental variation in damselfly body size, Ecography, 36: 894-903.

Mlynarek, J.M., Hassall, C. and Forbes, M.R. (2012) Higher gregarine parasitism often in sibling species of host damselflies with smaller geographical distributions, Ecological Entomology, 37: 419-425.

Hassall, C. (2012) Predicting the distributions of under-recorded Odonata using species distribution models, Insect Conservation and Diversity, 5: 192-201.

Penney, H.D., Hassall, C., Skevington, J.H., Abbott, K.R. and Sherratt, T.N. (2012) A comparative analysis of the evolution of imperfect mimicry. Nature, 483: 461-464.

Hassall, C., Hollinshead, J. and Hull, A. (2012) Temporal dynamics of aquatic communities and implications for pond conservation, Biodiversity and Conservation, 21 (3): 829-852.

Hassall, C. and Thompson, D.J. (2012) Study design and mark recapture estimates of dispersal: a case study with the endangered damselfly Coenagrion mercuriale, Journal of Insect Conservation, 16 (1): 111-120.

Hassall, C., Hollinshead, J. and Hull, A. (2011) Environmental correlates of plant and invertebrate species richness in ponds, Biodiversity and Conservation, 20: 3189-3222.

Thompson, D.J., Hassall, C., Lowe, C.D. and Watts, P.C. (2011) Field estimates of reproductive success in a model insect: behavioural surrogates are poor predictors of fitness, Ecology Letters, 14: 905-913.

Hassall, C. and T.N. Sherratt (2011). Statistical inference and spatial patterns in correlates of IQ.Intelligence, 39: 303-310.

Sherratt, T.N., Hassall, C., Laird, R.A., Thompson, D.J. and Cordero-Rivera, A. (2011) A comparative analysis of senescence in adult damselflies and dragonflies, Journal of Evolutionary Biology, 24: 810-822.

Sherratt, T.N., Laird, R.A., Hassall, C., Lowe, C.D., Harvey, I.F., Watts, P.C., Cordero-Rivera, A. and Thompson, D.J. (2010). Empirical evidence of senescence in adult damselflies (Odonata: Zygoptera). Journal of Animal Ecology, 79 (5), 1034-1044.

Hassall, C., and Thompson, D.J. (2010) Accounting for recorder effort in the detection of range shifts from historical data. Methods in Ecology and Evolution, 1 (4), 343-350.

Hassall, C., Lowe, C.D., Harvey, I.F., Watts, P.C. and Thompson, D.J. (2010) Phenology determines seasonal variation in ectoparasite loads in a natural insect population. Ecological Entomology, 35, 514-522.

Hassall, C., Thompson, D.J. and Harvey, I.F. (2010) The impact of climate-induced distributional changes on the validity of biological water quality metrics. Environmental Monitoring and Assessment, 160, 451-456.

Hassall, C., Thompson, D.J. and Harvey, I.F. (2009) Variation in morphology between core and marginal populations of three British damselflies. Aquatic Insects, 31 (3), 187-197.

Hassall, C. and Thompson D.J. (2009) Variation in the wingspot size and asymmetry of Banded Demoiselle Calopteryx splendens (Harris, 1792). Journal of the British Dragonfly Society, 25 (1), 7-15.

Hassall, C., Thompson, D.J. and Harvey, I.F. (2008) Latitudinal variation in morphology in two damselfly species with contrasting range dynamics. European Journal of Entomology, 105, 939-944.

Hassall, C. and Thompson, D.J. (2008) The impact of environmental warming on Odonata – a review. International Journal of Odonatology, 11 (2), 131-153.

Hassall, C., Thompson, D.J. and Harvey, I.F. (2008) Wings of Coenagrion puella vary in shape at the northern range margin (Odonata: Coenagrionidae). International Journal of Odonatology, 11 (1), 35-41.

Hassall, C., Thompson, D.J., French, G.C. and Harvey, I.F. (2007) Historical changes in the phenology of British Odonata are related to climate. Global Change Biology, 13, 933-941.



Hassall, C., Hebbern, C.A. & Centen, C.J. (2013) Climate Change Denial in the Classroom, Skeptical Inquirer, 37 (3).

Hassall, C. (2009) Environment Strategy and Action Plans: Progress Made (Research Paper), National Assembly for Wales, Cardiff.

Hassall, C. (2009) Dragonflies and man, Antenna, 33 (2), 98-99.

Hassall, C. (2007) Causes and consequences of body size variation in a damselfly. Bedfordshire Naturalist, 62 (2), 58-62.